C1 inlining checks that the declared "callee holder" is linked, which can result in a failure to inline in some situations:
1. invokedynamic, which relies on old code in ciBytecodeStream::get_declared_method_holder() that forces the holder to be java.lang.invoke.MethodHandle, even though the actual callee is a special generated method like java/lang/invoke/Invokers$Holder.linkToTargetMethod().
2. when the declared holder is not linked but the holder of the actual target method is linked